A. thaliana siliques produce high levels of benzoic acid (BA) that also include GSL compounds such as 3-benzoyloxypropylGSL (3BZO) and 4-benzoyloxybutylGSL (4BZO). T-DNA insertions that eliminate functional transcripts of At1g04580 (AAO4) cause decreases in the levels of total benzoic acid (BA) and in the levels of benzoylated GSLs in seeds (Ibdah et al. 2009)